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  1. Is the Y chromosome of Drosophila an evolved supernumerary chromosome?Johannes H. P. Hackstein, Ron Hochstenbach, Elisabeth Hauschteck-Jungen & Leo W. Beukeboom - 1996 - Bioessays 18 (4):317-323.
    The Y chromosomes of most Drosophila species are necessary for male fertility but they are not involved in sex determination. They have many puzzling properties that resemble the effects caused by B chromosomes. Classical genetic and molecular studies reveal substantial affinities between Y and B chromosomes and suggest that the Y chromosomes of Drosophila are not degenerated homologues of the X chromosomes, but rather that their Y chromosomes evolved as specialized supernumeraries similar to classical B chromosomes.
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