While there is ongoing debate about the existence of basic emotions and about their status as natural kinds, these debates usually carry on under the assumption that basic emotions are modular and therefore cannot account for behavioral variability in emotional situations. Moreover, both sides of the debate have assumed that these putative features of basic emotions distinguish them as products of evolution rather than products of culture and experience. I argue that these assumptions are unwarranted, that there is empirical evidence (...) against them, and that evolutionary theory itself should not lead us to expect that behavioral invariability and modularity mark the distinction between evolved emotions and higher cognitive emotions. I further suggest that claims about behavioral invariability and modularity have functioned as defeasible conjectures aimed at helping test basic emotion theory. Finally, I draw out the implications of these claims for debates about the existence of basic emotions in humans. (shrink)