Abstract
The philosophical practices undertaken in Plato's Academy remain, in the words of Cherniss, a 'riddle'. Yet surviving accounts of the views of the first two scholarchs of Plato's Academy after his death, Speusippus and Xenocrates, reveal a sophisticated engagement with their teacher's ideas concerning the pursuit of knowledge and the ends of philosophy. Speusippus and Xenocrates transform Plato's views on epistemology and happiness, and thereby help to lay the groundwork for the transformation of philosophy in the Hellenistic era.