Aristotle on exceptions to essences in biology

In Benedikt Strobel & Georg Wöhrle (eds.), Angewandte Epistemologie in antiker Philosophie und Wissenschaft, AKAN-Einzelschriften 11. Trier: Wissenschaftlicher Verlag Trier. pp. 69-92 (2016)
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Abstract
Exceptions are often cited as a counterargument against formal causation. Against this I argue that Aristotle explicitly allows for exceptions to essences in his biological writings, and that he has a means of explaining them through formal causation – though this means that he has to slightly elaborate on his general case theory from the Posterior Analytics, by supplementing it with a special case application in the biological writings. Specifically for Aristotle an essential predication need not be a universal predication. Rather an essential predication is where a property belongs essentially to a species or genus. This essential predication serves a causal and explanatory role, and is not dependent on a corresponding universal predication.
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