For Uexküll, biology is the science of the organization of living beings. In the context of Entwicklungsmechanik, he refers to Driesch’s and Spemann’s experiments on the development of embryonic germ cells to prove that self-differentiating processes constitute organisms as natural objects. Uexküll focuses on the theory of such self-differentiating processes or organizations. The notion of organization implies for him a “technique of nature” that is capable of structuring organic and inorganic material according to plans and rules. These plans and rules (...) are part of the overall order of the world. As preformed sign systems or codes, they determine and regulate the development and existence of individual animal subjects in their specific Umwelten. (shrink)

In biosemiotics, living beings are not conceived of as the passive result of anonymous selection pressures acted upon through the course of evolution. Rather, organisms are considered active participants that influence, shape and re-shape other organisms, the surrounding environment, and eventually also their own constitutional and functional integrity. The traditional Darwinian division between natural and sexual selection seems insufficient to encompass the richness of these processes, particularly in light of recent knowledge on communicational processes in the realm of life. Here, (...) we introduce the concepts of semiotic selection and semiotic co-option which in part represent a reinterpretation of classical biological terms and, at the same time, keep explanations sensitive to semiosic processes taking place in living nature. We introduce the term ‘semiotic selection’ to emphasize the fact that actions of different semiotic subjects (selectors) will produce qualitatively different selection pressures. Thereafter, ‘semiotic co-option’ explains how semiotic selection may shape appearance in animals through remodelling existing forms and relations. Considering the event of co-option followed by the process of semiotic selection enables us to describe the evolution of semantic organs. (shrink)

In this essay we argue for the possibility to describe the co-presence of species in a community as a consortium built by acoustic codes, using mainly the examples of bird choruses. In this particular case, the consortium is maintained via the sound-tope that different bird species create by singing in a chorus. More generally, the formation of acoustic codes as well as cohesive communicative systems (the consortia) can be seen as a result of plastic adaptational behaviour of the specimen who (...) can solve and avoid conflicts both with conspecifics and with other species in the vicinity. Thus, sign-relations appear to resolve potential conflicts, and as a foundation for symbiotic aggregations. The spatio-temporal configuration of consortia—their chronotope—includes several eco-fields as respective to different functions of the participating organisms. Biological study is combined with a semiotic approach that, as we suggest, should be more often used together to effectively describe ecological processes. (shrink)

Any biological species of biparental organisms necessarily includes, and is fundamentally dependent on, sign processes between individuals. In this case, the natural category of the species is based on family resemblances, which is why a species is not a natural kind. We describe the mechanism that generates the family resemblance. An individual recognition window and biparental reproduction almost suffice as conditions to produce species naturally. This is due to assortativity of mating which is not based on certain individual traits, but (...) on the difference between individuals. The biosemiotic model described here explains what holds a species together. It also implies that boundaries of a species are fundamentally fuzzy, and that character displacement occurs in case of sympatry. Speciation is a special case of discretisation that is an inevitable result of any communication system in work. The biosemiotic mechanism provides the conditions and communicative restrictions for the origin and persistence of diversity in the realm of living systems. (shrink)

The paper focuses on the semiotic principles of the organisation of ecosystems, attempting to find concepts that point to relations and not to elements. (1) Consortium (the term introduced by Johannes Reinke around 1873) can be defined as a group of organisms connected via (sign) relations, or groups of interspecific semiosic links in biocoenosis. The consortial relations include trophic and topic relations, both implying a recognition (identification) of the object by an organism involved (these, i.e., are sign relations). These relations (...) are ecologically inheritable. (2) Umwelt (the term introduced by Jakob von Uexküll around 1909) can be defined as a set of relations an organism has in an ecosystem (as in a semiosphere). The formation of an umwelt is dependent on the modelling system of the organism. (3) Biophony (the term introduced by Bernie Krause around 2000) denotes the coordination of inter- and intra-species relations in a soundscape of a biological community. This can be seen as a special case of Komposition as defined by Jakob and Thure von Uexküll. (4) Ecological code (as introduced, e.g. by Alexander Levich around 1977) can be defined as the set of (sign) relations (regular irreducible correspondences) characteristic to an entire ecosystem. We also mention the concepts of ecomones and coactones (introduced by Marcel Florkin in 1965) as the substances which are responsible for mediation of ecological inter-individual relations. All the relations as sign-relations evidently imply both a static or structuralist description (in terms of codes), and a processual description (in terms of semiosis carried on by interpretation). We conclude that all the above mentioned concepts can be viewed as conceptually connected and are suitable for semiotic description of biological communities. (shrink)

We examine the possibility of shifting the concept of choice to the centre of the semiotic theory of learning. Thus, we define sign process (meaning-making) through the concept of choice: semiosis is the process of making choices between simultaneously provided options. We define semiotic learning as leaving traces by choices, while these traces influence further choices. We term such traces of choices memory. Further modification of these traces (constraints) will be called habituation. Organic needs are homeostatic mechanisms coupled with choice-making. (...) Needs and habits result in motivatedness. Semiosis as choice-making can be seen as a complementary description of the Peircean triadic model of semiosis; however, this can fit also the models of meaning-making worked out in other shools of semiotics. We also provide a sketch for a joint typology of semiosis and learning. (shrink)

This paper recognizes a specific correspondence between biological evolution and technological development and on this basis tries to set up a semioticapproach to the evolutionary phenomenon of exaptation. To do this, the existence of a historical-structural and pragmatic analogy between organs and tools is shown, which in turn implies on a communicative ground the dissolution of some of their traditional distinctive att ributes. Finally, a philosophical-analytical approach to natural and cultural functions is applied to define three types of exaptations.