In The Origin of Species (1859) Darwin challenged many of the most deeply-held beliefs of the Western world. Arguing for a material, not divine, origin of species, he showed that new species are achieved by "natural selection." The Origin communicates the enthusiasm of original thinking in an open, descriptive style, and Darwin's emphasis on the value of diversity speaks more strongly now than ever. As well as a stimulating introduction and detailed notes, this edition offers a register of the many (...) writers referred to by Darwin in the text. (shrink)

In exploring the nature of psychological explanation, this book looks at how psychologists theorize about the human ability to calculate, to speak a language and the like. It shows how good theorizing explains or tries to explain such abilities as perception and cognition. It recasts the familiar explanations of "intelligence" and "cognitive capacity" as put forward by philosophers such as Fodor, Dennett, and others in terms of a theory of explanation that makes established doctrine more intelligible to professionals and their (...) students.In particular, the book shows that vestigial adherence to the positivists' D-N model has distorted the view of philosophers of science about what psychologists (and biologists) do and has masked the real nature of explanation. Major sections in the book cover Analysis and Subsumption; Functional Analysis; Understanding Cognitive Capacities; and Historical Reflections.Robert Cummins is Associate Professor of Philosophy at the University of Illinois, Chicago Circle. A Bradford Book. (shrink)

APA PsycNET abstract: This is the first volume of a two-volume work on Probability and Induction. Because the writer holds that probability logic is identical with inductive logic, this work is devoted to philosophical problems concerning the nature of probability and inductive reasoning. The author rejects a statistical frequency basis for probability in favor of a logical relation between two statements or propositions. Probability "is the degree of confirmation of a hypothesis (or conclusion) on the basis of some given evidence (...) (or premises)." Furthermore, all principles and theorems of inductive logic are analytic, and the entire system is to be constructed by means of symbolic logic and semantic methods. This means that the author confines himself to the formalistic procedures of word and symbol systems. The resulting sentence or language structures are presumed to separate off logic from all subjectivist or psychological elements. Despite the abstractionism, the claim is made that if an inductive probability system of logic can be constructed it will have its practical application in mathematical statistics, and in various sciences. 16-page bibliography. (PsycINFO Database Record (c) 2016 APA, all rights reserved). (shrink)

The Darwinian revolution supposed and imposed a much broader and more complex concept of environment than that which, until that moment, had been considered by most as part of natural history. Until Darwin, the environment of living beings had been regarded, almost exclusively, as the inanimate surroundings. This pre-Darwinian notion of environment included physicochemical and climatic variables: the living beings themselves were scarcely considered, or they were regarded just as food to be assimilated. In contrast, with Darwin, the influence of (...) living surroundings began to be regarded as having decisive importance for understanding the distribution, configuration, and viability of living beings. The relationships established among different living beings became the main and more significant features of the biological environment. This conceptual change was an exigency derived from the formulation and justification of the theory of natural selection. The idea of struggle for existence supposed by that theory requires that the biotic environment be considered critical in determining the viability of each living being. (shrink)

How questions and why questions interact in complex ways within biological practice. One of the most fruitful accounts to think about this relation is the widely known systemic approach, which has its origin mainly in the works of Robert Cummins (1975, 1983). As we will show, this approach effectively manages to capture much of what biologists do, especially in areas such as molecular biology, neuroscience and neuroethology. Our aim with this work is to discuss the metatheoretical status of the presuppositions (...) behind systemic analysis. We will hold that these assumptions are of an empirical nature, and that they can be reconstructed as parts of a theory. (shrink)

In this paper I defend an etiological theory of biological functions (according to which the proper function of a trait is the effect for which it was selected by natural selection) against three objections which have been influential. I argue, contrary to Millikan, that it is wrong to base our defense of the theory on a rejection of conceptual analysis, for conceptual analysis does have an important role in philosophy of science. I also argue that biology requires a normative notion (...) of a "proper function", and that a normative notion is not ahistorical. (shrink)

I defend the historical definition of "function" originally given in my Language, Thought and Other Biological Categories (1984a). The definition was not offered in the spirit of conceptual analysis but is more akin to a theoretical definition of "function". A major theme is that nonhistorical analyses of "function" fail to deal adequately with items that are not capable of performing their functions.

Biological functions are dispositions or effects a trait has which explain the recent maintenance of the trait under natural selection. This is the "modern history" approach to functions. The approach is historical because to ascribe a function is to make a claim about the past, but the relevant past is the recent past; modern history rather than ancient.

Este libro analiza y reconstruye una de las teorías científicas que más discusiones han provocado en el ámbito de la biología, de la filosofía y de la sociedad: la teoría de la selección natural. Esta teoría, que ocupa un lugar central en la biología evolutiva, se encuentra en el centro de la revolución darwiniana, uno de los cambios más radicales ocurridos en la historia de la ciencia y, sin dudas, uno de los que más consecuencias han tenido sobre la ciencia (...) en general, sobre la filosofía y sobre el mismo modo en que nos percibimos en el mundo. -/- Si bien se trata de una teoría que parece sencilla de comprender, ha sido especialmente elusiva a los intentos de reconstrucción, no hay acuerdo con respecto a su estructura ni a cuáles son sus conceptos fundamentales. En este trabajo, asumiendo el marco conceptual brindado por el estructuralismo metateórico, se propone una reconstrucción que permite no solo dar cuenta adecuadamente del modo en que la teoría funciona en la práctica científica actual, sino que, además, permite explicitar la importancia filosófica antes aludida. -/- Frente a los desacuerdos marcados en la filosofía de la biología, pero también en textos y manuales de los biólogos evolutivos, respecto del modo de presentación adecuado y la naturaleza de la teoría, la estrategia reconstructiva consiste en discutir la teoría en su origen, tal como apareció en los textos de uno de sus codescubridores, Charles Darwin. La intención del libro es, entonces, no solo explicitar la estructura de la teoría, sino, además, su lugar en el pensamiento de Darwin, en la revolución darwiniana y en la biología evolutiva. (shrink)

This paper reviews the debate on the notion of biological function and on functional explanation as this takes place in philosophy. It describes the different perspectives, issues, intuitions, theories and arguments that have emerged. The author shows that the debate has been too heavily influenced by the concerns of a naturalistic philosophy of mind and argues that in order to improve our understanding of biology the attention should be shifted from the study of intuitions to the study of the actual (...) practice of biological inquiry. (shrink)

This paper is concerned with reasonings that purport to explain why certain organisms have certain traits by showing that their actual design is better than contrasting designs. Biologists call such reasonings 'functional explanations'. To avoid confusion with other uses of that phrase, I call them 'design explanations'. This paper discusses the structure of design explanations and how they contribute to scientific understanding. Design explanations are contrastive and often compare real organisms to hypothetical organisms that cannot possibly exist. They are not (...) causal but appeal to functional dependencies between an organism's different traits. These explanations point out that because an organism has certain traits, it cannot be alive if the trait to be explained were replaced by a specified alternative. They can be understood from a mechanistic point of view as revealing the constraints on what mechanisms can be alive. (shrink)

The Nature of Selection is a straightforward, self-contained introduction to philosophical and biological problems in evolutionary theory. It presents a powerful analysis of the evolutionary concepts of natural selection, fitness, and adaptation and clarifies controversial issues concerning altruism, group selection, and the idea that organisms are survival machines built for the good of the genes that inhabit them. "Sober's is the answering philosophical voice, the voice of a first-rate philosopher and a knowledgeable student of contemporary evolutionary theory. His book merits (...) broad attention among both communities. It should also inspire others to continue the conversation."-Philip Kitcher, Nature "Elliott Sober has made extraordinarily important contributions to our understanding of biological problems in evolutionary biology and causality. The Nature of Selection is a major contribution to understanding epistemological problems in evolutionary theory. I predict that it will have a long lasting place in the literature."-Richard C. Lewontin. (shrink)

Darwin'suse of final cause accords with the Aristotelian idea of finalcauses as explanatory types – as opposed to mechanical causes, which arealways particulars. In Wright's consequence etiology, anadaptation is explained by particular events, namely, its past consequences;hence, that etiology is mechanistic at bottom. This justifies Ghiselin'scharge that such versions of teleology trivialize the subject, But a purelymechanistic explanation of an adaptation allows it to appear coincidental.Patterns of outcome, whether biological or thermodynamic, cannot be explainedbytracing causal chains, even were that possible. (...) They are explicanda of aspecialkind. The form of their explanation, in statistical mechanics or by naturalselection, is not captured by statistical variants of the covering-law model orrelated models of explanation. In them as in classical teleology, types ofoutcome are cited to explain why there are outcomes of those types. But onlywhen types are explanatory by being selected for, as inexplanations of animal and human behavior as well as in Darwin's theory ofnatural selection, but not in statistical mechanics, is the explanationteleological. Darwin's theory is nontrivially teleological. (shrink)

Teleological explanations in evolutionary biology, from Cuvier to the present (and into the future), depend on the metaphor of design for heuristic power and predictive fertility.

Contemporary analyses of biological function almost invariably advocate a naturalistic analysis, grounding biological functions in some feature of the mind-independent world. Many recent accounts suggest that no single analysis will be appropriate for all cases of use and that biological teleology should be split into several distinct categories. This paper argues that such accounts have paid too little attention to the way in which functional language is used, concentrating instead on the types of situation in which it is used. An (...) example of the role of teleology in science is examined and, on the basis of conclusions drawn from this, an alternative unifying analysis is proposed. It is suggested that, contrary to naturalistic accounts, teleology in biology carries no ontological commitment whatsoever to any class of mind-independent entities or properties. Instead, it is best regarded as a methodological device which is used to focus interest, formulate research perspectives and facilitate the structuring of certain questions or types of question that are pertinent in a given context of interest. (shrink)

RESUMEN Pocas polémicas en el ámbito de la filosofía de la biología han llevado tanto espacio como la que existen alrededor de la explicación funcional y del lenguaje funcional. Varios de los enfoques principales que han tratado de reconstruir este tipo de explicación y elucidar tal lenguaje, aunque fuertemente heterogéneos, comparten la idea de que el lenguaje funcional no es más que un modo de hablar del cual puede prescindirse, pues, asumen que el lenguaje funcional puede ser definido a partir (...) de (o reducido a) lenguaje no funcional. En consecuencia, la explicación funcional sería reducible a (o equivalente a) explicaciones no funcionales. En este trabajo repasaremos tales enfoques, y trataremos de defender que no funcionan en el siguiente sentido: no pueden dar cuenta de la totalidad de las atribuciones funcionales. En consecuencia, aunque pudieran ser útiles para elucidar o reconstruir algunas áreas de la biología, no cumplen con el objetivo de mostrar que el lenguaje funcional es eliminable. -/- RESUMO Poucas polêmicas no âmbito da filosofia da biologia tiveram tanto espaço como a que existe acerca da explicação funcional e da linguagem funcional. Vários dos principais enfoques que trataram de reconstruir este tipo de explicação e elucidar tal linguagem, ainda que fortemente heterogêneos, compartilham a ideia de que a linguagem funcional não é mais que um modo de falar do qual pode prescindir-se, pois, assumem que a linguagem funcional pode ser definida a partir de (ou reduzida a) linguagem não funcional. Em consequência, a explicação funcional seria reduzível a (ou equivalente a) explicações não funcionais. Neste trabalho repassaremos tais enfoques e trataremos de defender que não funcionam, no seguinte sentido: não podem dar conta da totalidade das atribuições funcionais. Em consequência, ainda que pudessem ser úteis para elucidar ou reconstruir algumas áreas da biologia, não alcançam o objetivo de mostrar que a linguagem funcional é eliminável. (shrink)

Functional language is ubiquitous in ecology, mainly in the researches about biodiversity and ecosystem function. However, it has not been adequately investigated by ecologists or philosophers of ecology. In the contemporary philosophy of ecology we can recognize a kind of implicit consensus about this issue: while the etiological approaches cannot offer a good concept of function in ecology, Cummins’ systemic approach can. Here we propose to go beyond this implicit consensus, because we think these approaches are not adequate for ecology. (...) We argue that a sound epistemological framework to function in ecology is to be found in organizational approaches. In this line, we define function in ecology as a precise effect of a given constraint on the ecosystem flow of matter and energy performed by a given item of biodiversity, within a closure of constraints. We elaborate on this definition by developing a case study of a bromeliad ecosystem. (shrink)

This paper argues that a minimal notion of function and a notion of normal-proper function are used in explaining how bodies and brains operate. Neither is Cummins’ notion, as originally defined, and yet his is often taken to be the clearly relevant notion for such an explanatory context. This paper also explains how adverting to normal-proper functions, even if these are selected functions, can play a significant scientific role in the operational explanations of complex systems that physiologists and neurophysiologists provide, (...) despite a lack of relevant causal efficacy on the part of such functions. (shrink)

In this paper, we develop an organizational account that defines biological functions as causal relations subject to closure in living systems, interpreted as the most typical example of organizationally closed and differentiated self-maintaining systems. We argue that this account adequately grounds the teleological and normative dimensions of functions in the current organization of a system, insofar as it provides an explanation for the existence of the function bearer and, at the same time, identifies in a non-arbitrary way the norms that (...) functions are supposed to obey. Accordingly, we suggest that the organizational account combines the etiological and dispositional perspectives in an integrated theoretical framework. IntroductionDispositional ApproachesEtiological TheoriesBiological Self-maintenance Closure, teleology, and normativityOrganizational differentiationFunctions C1: Contributing to the maintenance of the organization C2: Producing the functional trait Implications and Objections Functional versus useful Dysfunctions, side effects, and accidental contributionsProper functions and selected effectsReproductionRelation with other ‘unitarian’ approachesConclusions. (shrink)

The concept of mechanism is analyzed in terms of entities and activities, organized such that they are productive of regular changes. Examples show how mechanisms work in neurobiology and molecular biology. Thinking in terms of mechanisms provides a new framework for addressing many traditional philosophical issues: causality, laws, explanation, reduction, and scientific change.

It is often claimed that one of Darwin''s chief accomplishments was to provide biology with a non-teleological explanation of adaptation. A number of Darwin''s closest associates, however, and Darwin himself, did not see it that way. In order to assess whether Darwin''s version of evolutionary theory does or does not employ teleological explanation, two of his botanical studies are examined. The result of this examination is that Darwin sees selection explanations of adaptations as teleological explanations. The confusion in the nineteenth (...) century about Darwin''s attitude to teleology is argued to be a result of Darwin''s teleological explanations not conforming to either of the dominant philosophical justifications of teleology at that time. Darwin''s explanatory practices conform well, however, to recent defenses of the teleological character of selection explanations. (shrink)

Aristotle’s principle that “nature does nothing in vain” (NDNIV) is central to his teleological approach to understanding organisms. First, we argue that James G. Lennox’s influential account of NDNIV is unsuccessful. Second, we propose an alternative account that includes a natural state model. According to a natural state model of development, an organism will develop toward its natural state unless interfering forces prevent that from happening. Third, we argue that this account also fits Aristotle’s discussion in the Generation of Animals (...) and can explain monstrosities in nature that at first sight seem to be counterexamples to NDNIV. Fourth, we take a broader look at NDNIV, arguing that it does not entail that all structures are teleological (since it accommodates the thought that by-products and some neglected items that Aristotle calls “tokens” are nonteleological), that it does not entail that there is only one function per organ, that NDNIV is not an anthropocentric principle, and that, perhaps surprisingly, the teleological structures of an organism can benefit others apart from the organism and its progeny. Finally, we argue that Aristotle endorsed an empirical justification of NDNIV wherein the principle is justified by its many successful applications. (shrink)

Suele señalarse la fuerte influencia que la teología natural tuvo sobre Darwin en su conceptualización del fenómeno de la adaptación. La teoría de la selección natural explicaría el mismo fenómeno que los teólogos naturales querían explicar: la adaptación. Recientemente ha sido señalado, sin embargo, que la forma darwiniana de conceptualizar la adaptación es novedosa. Las adaptaciones de la teología natural presupondrían la idea de que los organismos existen para la realización y manutención del equilibrio natural establecido por el creador. El (...) punto de este trabajo discutir esta cuestión, asumiendo el marco del estructuralismo metateórico. The strong influence that natural theology had over Darwin’s conceptualization of the adaptation phenomenon is frequently noted. The natural selection theory would explain the same phenomenon that the natural theologist intended to explain: the adaptation. Recently has been claimed, on the contrary, that the Darwinian way of conceptualizing adaptation is new. The concept of adaptation from natural theology presupposes the idea that organisms exist for realization and support of the natural equilibrium stated by the creator. In this work I will discuss this point, assuming the metatheoretical structuralism frame. (shrink)

The concept of fitness has generated a lot of discussion in philosophy of biology. There is, however, relative agreement about the need to distinguish at least two uses of the term: ecological fitness on the one hand, and population genetics fitness on the other. The goal of this paper is to give an explication of the concept of ecological fitness by providing a reconstruction of the theory of natural selection in which this concept was framed, that is, based on the (...) way the theory was put to use in Darwin’s main texts. I will contend that this reconstruction enables us to account for the current use of the theory of natural selection. The framework presupposed in the analysis will be that of metatheoretical structuralism. This framework will provide both a better understanding of the nature of ecological fitness and a more complete reconstruction of the theory. In particular, it will provide what I think is a better way of understanding how the concept of fitness is applied through heterogeneous cases. One of the major advantages of my way of thinking about natural selection theory is that it would not have the peculiar metatheoretical status that it has in other available views. I will argue that in order to achieve these goals it is necessary to make several concepts explicit, concepts that are frequently omitted in usual reconstructions. (shrink)

Darwin''s biology was teleological only if the term teleology is defined in a manner that fails to recognize his contribution to the metaphysics and epistemology of modern science. His use of teleological metaphors in a strictly teleonomic context is irrelevant to the meaning of his discourse. The myth of Darwin''s alleged teleology is partly due to misinterpretations of discussions about whether morphology should be a purely formal science. Merely rejecting such notions as special creation and vitalism does not prevent the (...) pernicious effects of teleological reasoning, even at the present time. (shrink)

I present and defend the generalized selected effects theory (GSE) of function. According to GSE, the function of a trait consists in the activity that contributed to its bearer’s differential reproduction, or differential retention, within a population. Unlike the traditional selected effects (SE) theory, it does not require that the functional trait helped its bearer reproduce; differential retention is enough. Although the core theory has been presented previously, I go significantly beyond those presentations by providing a new argument for GSE (...) and defending it from a recent objection. I also sketch its implications for teleosemantics and philosophy of medicine. (shrink)

The purpose of this paper is to defend, contra Fodor and Piattelli-Palmarini (F&PP), that the theory of natural selection (NS) is a perfectly bona fide empirical unified explanatory theory. F&PP claim there is nothing non-truistic, counterfactual-supporting, of an “adaptive” character and common to different explanations of trait evolution. In his debate with Fodor, and in other works, Sober defends NS but claims that, compared with classical mechanics (CM) and other standard theories, NS is peculiar in that its explanatory models are (...) a priori (a trait shared with few other theories). We argue that NS provides perfectly bona fide adaptive explanations of phenotype evolution, unified by a common natural-selection guiding principle. First, we introduce the debate and reply to F&PP’s main argument against NS. Then, by reviewing different examples and analyzing Fisher’s model in detail, we show that NS explanations of phenotypic evolution share a General Natural Selection Principle. Third, by elaborating an analogy with CM, we argue against F&PP’s claim that such a principle would be a mere truism and thus explanatorily useless, and against Sober’s thesis that NS models/explanations have a priori components that are not present in CM and other common empirical theories. Irrespective of differences in other respects, the NS guiding principle has the same epistemic status as other guiding principles in other highly unified theories such as CM. We argue that only by pointing to the guiding principle-driven nature that it shares with CM and other highly unified theories, something no-one has done yet in this debate, one can definitively show that NS is not defective in F&PP’s sense: in the respects relevant to the debate, Natural Selection is as defective and as epistemically peculiar as Classical Mechanics and other never questioned theories. (shrink)

The epistemic status of Natural Selection has seemed intriguing to biologists and philosophers since the very beginning of the theory to our present times. One prominent contemporary example is Elliott Sober, who claims that NS, and some other theories in biology, and maybe in economics, are peculiar in including explanatory models/conditionals that are a priori in a sense in which explanatory models/conditionals in Classical Mechanics and most other standard theories are not. Sober’s argument focuses on some “would promote” sentences that (...) according to him, play a central role in NS explanations and are both causal and a priori. Lange and Rosenberg criticize Sober arguing that, though there may be some unspecific a priori causal claims, there are not a priori causal claims that specify particular causal factors. Although we basically agree with Lange and Rosenberg’s criticism, we think it remains silent about a second important element in Sober’s dialectics, namely his claim that, contrary to what happens in mechanics, in NS explanatory conditionals are a priori, and that this is so in quite specific explanatory models. In this paper we criticize this second element of Sober’s argument by analyzing what we take to be the four possible interpretations of Sober’s claim, and argue that, terminological preferences aside, the possible senses in which explanatory models in NS can qualify, or include elements that can qualify, as a priori, also apply to CM and other standard, highly unified theories. We conclude that this second claim is unsound, or at least that more needs to be said in order to sustain that NS explanatory models are a priori in a sense in which CM models are not. (shrink)

A coherent treatment of the flow of ideas throughout Darwin's works, this volume presents a unified theoretical system that explains Darwin's investigations, evaluating the literature from a historical, scientific, and philosophical perspective.

Explores the development of the ideas of evolutionary biology, particularly as affected by the increasing understanding of genetics and of the chemical basis of inheritance.

Carl F. Craver investigates what we are doing when we use neuroscience to explain what's going on in the brain. When does an explanation succeed and when does it fail? Craver offers explicit standards for successful explanation of the workings of the brain, on the basis of a systematic view about what neuroscientific explanations are.

Provides a philosophical analysis of such biological concepts as natural selection, adaptation, speciation, and evolution.

FEW HEROES LOWER their sights in the prime of their lives; triumph leads inexorably on, often to destruction. Alexander wept because he had no new worlds to conquer; Napoleon, overextended, sealed his doom in the depth of a Russian winter. But Charles Darwin did not follow the Origin of Species (1859) with a general defense of natural selection or with its evident extension to human evolution (he waited until 1871 to publish The Descent of Man). Instead, he wrote his most (...) obscure work, a book entitled: On the Various Contrivances by Which British and Foreign Orchids Are Fertilized by Insects (1862). (shrink)

En la Teoría de la Selección Natural, el concepto de función biológica debe suponerse para delimitar el concepto de aptitud; y éste debe suponerse para delimitar el concepto de adaptación y también para explicar el fenómeno al que este último alude. Esos tres conceptos, por otra parte, son especificaciones de tres conceptos de aplicación más universal. El concepto de función biológica es un caso particular del concepto general de función; y el concepto de aptitud especifica el concepto de eficiencia. El (...) concepto de adaptación, mientras tanto, es una especificación del concepto de diseño. La visualización de esas articulaciones conceptuales permite entender la legitimidad de la naturalización de la teleología operada por el darwinismo.In Natural Selection Theory, the concept of biological function must be supposed to elucidate the concept of fitness; and this one must be supposed to elucidate the concept of adaptation and also to explain the phenomenon to which the last one alludes. These three concepts, on the other hand, are specificationsof three concepts of more universal application. The concept of biological function is a particular case of the general concept of function; and the concept of fitness is a specification of the concept of efficiency. The concept of adaptation, meanwhile, specifies the concept of design. The visualization of these conceptualjoints allows to understanding the legitimacy of the naturalization of the teleology produced by the Darwinism. (shrink)