A study is made of the history of the type and related concepts, from Greek Antiquity up to the present. It is demonstrated that the type-concept of eighteenth century biology was based on Leibniz's concept of substantial form, and was not related to a Platonic Idea, whilst it is now generally understood in the sense of model or norm. In the present paper, a type-concept is developed which includes ontogenetic and phylogenetic time and various evolutionary mechanisms. This type can serve (...) as a model of the evolutionary potentialities of a taxon, and as a standard of higher classification. All classifications based on the same archetype, whether typological, numerical or phylogenetic, will be comparable, although not necessarily identical. (shrink)

The sole diagram in On the Origin of Species is generally considered to be merely an illustration of Darwin’s ideas, but such an interpretation ignores the fact that Darwin himself expressly stated that the diagram helped him to discover and express his ideas. This article demonstrates that developing the so-called “tree diagram” substantially aided Darwin’s heuristics. This demonstration is based on an interpretation of the diagram and of 17 sketches found in Darwin’s scientific papers. The key to this interpretation is (...) the meaning that Darwin assigned to the graphic elements he used to construct the preliminary sketches and the diagram. I argue that each of the sketches contributed to the shaping of Darwin’s ideas and that, in their succession, each added new elements that ultimately resulted in the fully developed published diagram. (shrink)

Information plays an important role not only in evolution—genetics can be seen as a mechanism to transfer information from one generation to the next—but also in ecology: virtually all organisms use information about their environment to adjust their behavior and life histories. Indeed, being adapted to something can be defined as having the right information to solve the life-history problems that this creates. It then becomes irrelevant precisely where this information came from (genetics, experience, culture, etc.) but rather becomes an (...) issue of how to assess the fitness value of this information. Here I will explore in some detail the conceptual consequences of the information-based view of evolution. In particular, I will focus on how individuals are actually associations of smaller units that depend on each other’s genetic information and on information from multiple external sources. An important and fundamental problem arises when these different sources of information do not suggest the same adaptations. Conflicting information may thus give new insight into the causes of stress. (shrink)

Opatrzność Boża a przypadek w świecie Celem artykułu jest obrona dwóch tez: pierwszej, że istnienie zdarzeń przypadkowych jest do pogodzenia z istnieniem Boga oraz tezy drugiej, że przypadek może być częścią Bożej opatrzności. Koniunkcja obu powyższych tez nazwana jest w artykule tezą kompatybilizmu. Argumentacja w obronie kompatybilizmu opiera się na danych współczesnej nauki oraz na idei wszechmocnego Boga Stwórcy. Porządek argumentacji w artykule jest następujący. W części drugiej przedstawiony jest historyczny kontekst oraz podstawy doktrynalne pojęcia opatrzności. W części trzeciej omówiony (...) jest argument za nieistnieniem Boga oparty na założeniu głoszącym istnienie zdarzeń przypadkowych w świecie oraz przedstawiona jest krytyka tego argumentu. W części czwartej prezentowane są możliwe modele działania Boga w świecie, w ramach których przyjmuje się istnienie zdarzeń przypadkowych. W części piątej i ostatniej rozważana jest idea Bożej doskonałości obejmującej takie atrybuty jak wszechmoc, wszechwiedzę, doskonałą dobroć i suwerenną wolę Bożą. (shrink)

Philosophers agree that scientific explanations aim to produce understanding, and that good ones succeed in this aim. But few seriously consider what understanding is, or what the cues are when we have it. If it is a psychological state or process, describing its specific nature is the job of psychological theorizing. This article examines the role of understanding in scientific explanation. It warns that the seductive, phenomenological sense of understanding is often, but mistakenly, viewed as a cue of genuine understanding. (...) The article closes with a discussion of several new paths of research that tie the psychology of scientific explanation to cognate notions of learning, testimony, and understanding. (shrink)

In a macroevolutionary timescale, evolvability itself evolves. Lineages are sorted based on their ability to generate adaptive novelties, which leads to the optimization of their genotype-phenotype map. The system of translation of genetic or epigenetic changes to the phenotype may reach significant horizontal and vertical complexity, and may even exhibit certain aspects of learning behaviour. This continuously evolving semiotic system probably enables the origin of complex yet functional and internally compatible adaptations. However, it also has a second, “darker”, side. As (...) was pointed out by several authors, the same process gradually reduces the probability of the origination of significant evolutionary novelties. In a similar way to the evolution of societies, teachings, or languages, in which the growing number of internal linkages gradually solidifies their overall structure and the structure or interpretation of their constitutive elements, the evolutionary potential of lineages decreases during biological evolution. Possible adaptations become limited to small “peripheral” modifications. According to the Frozen Evolution theory, some of the proximate causes of this “macroevolutionary freezing” are more pronounced or present exclusively in sexual lineages. Sorting based on the highest evolvability probably leads to the establishment of certain structural features of complex organisms, e.g. the modular character of their development and morphology. However, modules also “macroevolutionary freeze” whereas the hypothetical “thawing” of modules or their novel adaptive combinations becomes rarer and rarer. Some possible ways out of this dead end include the rearrangement of individual development, e.g. neoteny, radical simplification, i.e. sacculinization, and transition to a higher level of organization, e.g. symbiosis or symbiogenesis. The evolution of evolvability is essentially a biosemiotic process situated at the intersection of the genocentric modern synthesis and the evo-devo-centric extended synthesis. Therefore, evolvability may eventually connect these three not necessarily contradictory approaches. (shrink)

Super-substantivalism is the thesis that space is identical to matter; it is currently under discussion ? see Sklar (1977, 221?4), Earman (1989, 115?6) and Schaffer (2009) ? in contemporary philosophy of physics and metaphysics. Given this current interest, it is worth investigating the thesis in the history of philosophy. This paper examines the super-substantivalism of Samuel Alexander, an early twentieth century metaphysician primarily associated with (the movement now known as) British Emergentism. Alexander argues that spacetime is ontologically fundamental and it (...) gives rise to an ontological hierarchy of emergence, involving novel properties such as matter, life and mind. Alexander's super-substantivalism is interesting not just because of its historical importance but also because Alexander unusually attempts to explain why spacetime is identical to matter. This paper carefully unpacks that explanation and shows how Alexander is best read as conceiving of spacetime as a Spinozistic substance, worked upon by evolution. (shrink)

Song in oscine birds relies upon the rare capacity of vocal learning. Transmission can be vertical, horizontal, or oblique. As a rule, memorization and production by a naïve bird are not simultaneous: the long-term storage of song phrases precedes their first vocal rehearsal by months. While a wealth of detail regarding songbird enculturation has been uncovered by focusing on the apprentice, whether observational learning can fully account for the ontogeny of birdsong, or whether there could also be an element of (...) active teaching involved, has remained an open question. Given the paucity of knowledge on animal cultures, I argue for the utility of an inclusive definition of teaching that encourages data be collected across a wide range of taxa. Borrowing insights from musicology, I introduce the Australian pied butcherbird into the debate surrounding mechanisms of cultural transmission. I probe the relevance and utility of mentalistic, culture-based, and functionalist approaches to teaching in this species. Sonographic analysis of birdsong recordings and observational data of pied butcherbird behavior at one field site provide evidence that I assess based on criteria laid down by Caro and Hauser, along with later refinements to their functionalist definition. The candidate case of teaching reviewed here adds to a limited but growing body of reports supporting the notion that teaching may be more widespread than is currently realized. Nonetheless, I describe the challenges of confirming that learning has occurred in songbird pupils, given the delay between vocal instruction and production, as well as the low status accorded to anecdote and other observational evidence commonly mustered in instances of purported teaching. As a corrective, I press for an emphasis on biodiversity that will guide the study of teaching beyond human accounts and intractable discipline-specific burdens of proof. (shrink)

Welch :263–279, 2017) has recently proposed two possible explanations for why the field of evolutionary biology is plagued by a steady stream of claims that it needs urgent reform. It is either seriously deficient and incapable of incorporating ideas that are new, relevant and plausible or it is not seriously deficient at all but is prone to attracting discontent and to the championing of ideas that are not very relevant, plausible and/or not really new. He argues for the second explanation. (...) This paper presents a twofold critique of his analysis: firstly, the main calls for reform do not concern the field of evolutionary biology in general but rather, or more specifically, the modern evolutionary synthesis. Secondly, and most importantly, these calls are not only inspired by the factors, enumerated by Welch, but are also, and even primarily, motivated by four problematic characteristics of the modern synthesis. This point is illustrated through a short analysis of the latest reform challenge to the modern synthesis, the so-called extended evolutionary synthesis. We conclude with the suggestion that the modern synthesis should be amended, rather than replaced. (shrink)

Traditionally, Thomas S. Kuhn’s The Structure of Scientific Revolutions (1962) is largely identified with his analysis of the structure of scientific revolutions. Here, we contribute to a minority tradition in the Kuhn literature by interpreting the history of evolutionary biology through the prism of the entire historical developmental model of sciences that he elaborates in The Structure. This research not only reveals a certain match between this model and the history of evolutionary biology but, more importantly, also sheds new light (...) on several episodes in that history, and particularly on the publication of Charles Darwin’s On the Origin of Species (1859), the construction of the modern evolutionary synthesis, the chronic discontent with it, and the latest expression of that discontent, called the extended evolutionary synthesis. Lastly, we also explain why this kind of analysis hasn’t been done before. (shrink)

It has long been taken for granted that the immune system’s capacity to protect an individual from infection and disease depends on the power of the system to distinguish between self and nonself. However, accumulating data have undermined this fundamental concept. Evidence against the self/nonself discrimination model left researchers in need of a new overarching framework able to capture the immune system’s reactivity. Here, I highlight that along with the self/nonself model, another powerful representation of the immune system’s reactivity has (...) been developed in the twentieth century immunology. According to this alternative view, the immune system is not a killer of nonself strangers but a peace-maker helping to establish harmony with the environment. The balance view of the system has never become part of the dominant paradigm. However, it is gaining more and more currency as new research develops. Advances in mucosal immunology confirm that instead of distinguishing between self and foreign the immune system reacts to microbial, chemical and self-induced alterations to produce responses that counterbalance effects of these changes. (shrink)

The evolutionary origin of consciousness has been a growing area of study in recent years. Nevertheless, there is intense debate on whether the existence of phenomenal consciousness without the cerebral cortex is possible. The corticocentrists have an empirical advantage because we are quasi-confident that we humans are conscious and have the well-developed cortex as the site of our consciousness. However, their prejudice can be an anthropic bias similar to the anthropocentric prejudice in pre-Darwinian natural history. In this paper, I propose (...) three basic principles to provide a conceptual basis for evolutionary studies of consciousness: the non-solipsistic principle, the evolution principle, and the anthropic principle. These principles collectively help us to avoid solipsism, anthropocentrism, and anthropomorphism to some degree, although we cannot be completely free from them. Also, the landscape metaphor associated with the anthropic principle provides an image of how different forms of consciousness can be acquired. (shrink)

This paper attempts to reconcile the conceptual conflict between typological and population thinking to provide a philosophical foundation for extended evolutionary synthesis. Typological thinking has been considered a pre-Darwinian, essentialist dogma incompatible with population thinking, which is the core notion of Darwinism. More recent philosophical and historical studies suggest that a non-essentialist form of typology has some advantages in the study of evolutionary biology. However, even if we adopt such an epistemological interpretation of typological thinking, there still remains an epistemological (...) and methodological conflict between these two styles of thinking. How can we relate typological thinking with population thinking in pursuit of more integrated or interconnected research into evolutionary biology? I propose that homology thinking, which is another style of thinking that recognizes homologous characters, provides a common basis for typological representations of character states and for character dynamics in an evolving population. Good examples of this bridging role are found in teratology and breeding, where variation and novelty are recognized in developmental and morphological traits, gene expression patterns, and so on. Essentialism-free, dynamic views of homology have great potential to reconcile typological and population thinking and to set the stage for the EES. (shrink)

Evolutionary adaptation has been suggested as the hallmark of life that best accounts for life’s creativity. However, current evolutionary approaches still fail to give an adequate account of it, even if they are able to explain both the origin of novelties and the proliferation of certain traits in a population. Although modern-synthesis Darwinism is today usually appraised as too narrow a position to cope with all the complexities of developmental and structural biology—not to say biosemiotic phenomena—, Darwinism need not be (...) if we separate metaphor from reality in natural selection in order to show the axiological complexity of this concept. This can shed light on the relationship between biosemiotics and biological evolution. (shrink)

If there is a single discipline of science calling the basic concepts of biology into question, it is without doubt microbiology. Indeed, developments in microbiology have recently forced us to rethink such fundamental concepts as the organism, individual, and genome. In this paper I show how microorganisms are changing our understanding of natural aggregations and develop the concept of a Darwinian population to embrace these discoveries. I start by showing that it is hard to set the boundaries of a Darwinian (...) population, and I suggest thinking of a Darwinian population as a relative property of a Darwinian individual. Then I argue, in contrast to the commonly held view, that Darwinian populations are multispecies units, and that in order to accept the multispecies account of Darwinian populations we have to separate fitness from natural selection. Finally, I show how all these ideas provide a theoretical framework leading to a more precise understanding of the ecology of endosymbiosis than is afforded by poetic metaphors such as ‘slavery’. (shrink)

Leigh Van Valen was an American evolutionary biologist who made major contributions to evolutionary theory. He is particularly remembered for his groundbreaking paper “A New Evolutionary Law” (1973) where he provided evidence from fossil record data that the probability of extinction within any group remains essentially constant through time. In order to explain such an unexpected result, Van Valen formulated a very influential idea that he dubbed the “Red Queen hypothesis.” It states that the constant decay must be a consequence (...) of evolutionary interactions among connected species within ecological networks. In Van Valen’s picture, species do not merely evolve: they also coevolve with other species. As a consequence, when thinking about adaptation to an external environment, the other species must be considered as part (maybe a major part) of such an external world. Van Valen’s law provided the first complex systems theory of coevolutionary dynamics and inspired a whole range of theoretical and experimental developments from very diverse fields, percolating far beyond its original formulation. Red Queen arms races are nowadays considered a widespread feature of complex adaptive systems. (shrink)

This book consists of four essays on Darwin’s theory of evolution and three postscripts on contemporary evolutionary theory. The main questions that the chapters address are: How are common ancestry and natural selection related to each other in Darwin’s theory? What were Darwin’s views about group selection and the evolution of altruism? Why did Darwin change his mind about sex ratio, and how are his ideas related to the creationism that preceded him and the evolutionary biology that followed? Do Darwin’s (...) theory and modern evolutionary theory obey the principle of methodological naturalism and ought they to do so?The book is philosophical because it focuses on the logic of arguments. I attend closely to Darwin’s texts, but I sometimes use tools that Darwin did not have available. These include ideas from contemporary biology and from probability theory. I see merit in this approach because it helps elucidate Darwin’s theory. Darwin pointed to the need for this sort of .. (shrink)

The reception so far of Kitcher's Vaulting Ambition reminds me of the old saw about the difference between an optimist and a pessimist. Looking at the same glass of water, the former sees it as half full while the latter sees it as half empty. Some have seen Kitcher's book as a vindication of the possibility of an evolutionary science of human behavior; others have seen it as a devastating critique of the most influential efforts to date to construct such (...) a science. As in the joke about the water glass, both assessments have their point. Sociobiology is a research program, not a unified theory. What one can do is test particular hypotheses within this program and see whether they live up to reasonable standards of rigor and evidence. This is what Kitcher does, with great energy and care. (shrink)

To evaluate Hume's thesis that causal claims are always empirical, I consider three kinds of causal statement: ?e1 caused e2 ?, ?e1 promoted e2 ?, and ?e1 would promote e2 ?. Restricting my attention to cases in which ?e1 occurred? and ?e2 occurred? are both empirical, I argue that Hume was right about the first two, but wrong about the third. Standard causal models of natural selection that have this third form are a priori mathematical truths. Some are obvious, others (...) less so. Empirical work on natural selection takes the form of defending causal claims of the first two types. I provide biological examples that illustrate differences among these three kinds of causal claim. (shrink)

This article argues that feminist analyses of patriarchy should be expanded to address the evolutionary basis of male motivation to control female sexuality. Evidence from other primates of male sexual coercion and female resistance to it indicates that the sexual conflicts of interest that underlie patriarchy predate the emergence of the human species. Humans, however, exhibit more extensive male dominance and male control of female sexuality than is shown by most other primates. Six hypotheses are proposed to explain how, over (...) the course of human evolution, this unusual degree of gender inequality came about. This approach emphasizes behavioral flexibility, cross-cultural variability in the degree of partriarchy, and possibilities for future change. (shrink)

An increasing number of biologists are expressing discontent with the prevailing theory of neo-Darwinism. In particular, the tendency of neo-Darwinians to adopt genetic determinism and atomistic notions of both genes and organisms is seen as grossly unfair to the body of developmental theory. One faction of dissenteers, the Process Structuralists, take their inspiration from the rational morphologists who preceded Darwin. These neo-rationalists argue that a mature biology must possess universal laws and that these generative laws should be sought within organismal (...) development. Such a rational biology will only be possible once the neo-Darwinian paradigm, with its reliance on inherently stochastic processes, is overthrown.To facilitate this revolution, process structuralism launches a broad attack on the theoritical adequacy of its opponent. It is charged that neo-Darwinism is untestable and therefore its hypotheses are nothing more than adaptive stories. Further, the lamentable tendencies toward genetic determinism and atomism by modern biologists is seen as the inescapable consequences of adopting the neo-Darwinian outlook. (shrink)

Nietzsche was a philosopher, not a biologist, Nevertheless his philosophical thought was deeply influenced by ideas emerging from the evolutionary biology of the nineteenth century. His relationship to the Darwinism of his time is difficult to disentangle. It is argued that he was in a sense an unwitting Darwinist. It follows that his philosophical thought is of considerable interest to those concerned to develop an evolutionary biology of mankind. His approach can be likened to that of an extraterrestrial sociobiologist studying (...) clever beasts... in some out of the way corner of the universe ... It is shown how be uses this viewpoint to account for the origin of the central psychobiology of humankind: for dualistic philosophies, such as that of Descartes (which Ryle famously called the official doctrine), for human notions of truth and falsehood, being and becoming, and for other fundamental concepts of Western philosophy and science. All these, he argues, are no more and no less than the necessary adaptations of a zoological species, Homo sapiens, in its struggle for life in a Darwinian world. It is concluded that Nietzsche was the first philosopher to accept and use in their full depth the philosophical implications of nineteeth-century evolutionism, implications which are still resisted to this day. It is also argued that this interpretation of Nietzsche's aphoristic writings provides them with an organic consistency. (shrink)

Two prominent theoretical frameworks in moral psychology, Moral Foundations and Dual Process Theory, share a broad foundational assumption that individual differences in human morality are dispositional and in part due to genetic variation. The only published direct test of heritability, however, found little evidence of genetic influences on moral judgments using instrumentation approaches associated with Moral Foundations Theory. This raised questions about one of the core assumptions underpinning intuitionist theories of moral psychology. Here we examine the heritability of moral psychology (...) using the moral dilemmas approach commonly used in Dual Process Theory research. Using such measures, we find consistent and significant evidence of heritability. These findings have important implications not only for understanding which measures do, or do not, tap into the genetically influenced aspects of moral decision-making, but in better establishing the utility and validity of different intuitionist theoretical frameworks and the source of why people differ in those frameworks. (shrink)

Understanding female sexuality and mate choice is central to evolutionary scenarios of human social systems. Studies of female sexuality conducted by sex researchers in the United States since 1938 indicate that human females in general are concerned with their sexual well-being and are capable of sexual response parallel to that of males. Across cultures in general and in western societies in particular, females engage in extramarital affairs regularly, regardless of punishment by males or social disapproval. Families are usually concerned with (...) marriage arrangements only insofar as those arrangements are economically or politically advantageous, but females most often have a voice in arranged marriages. Extended families also concentrate on a couple’s future reproduction rather than on sexual exclusivity. Although marriage for females is often compromised by male or family reproductive interests (which may not in fact differ from female interests), females appear to exercise their sexuality with more freedom than has previously been suggested. Notions of human females as pawns in the male reproductive game, or as traders of sex for male services, should be dispelled. (shrink)

The relationship between the Bible and science has been debated for decades. While science has emerged as a multifaceted discipline focused on the natural world, it has been viewed as a growing body of facts or knowledge ; and a path to understanding. As scientists test ideas, emerging disciplines such as palaeoanthropology, geology, archaeology, and evolutionary biology have attempted to prove Christian beliefs based on the Biblical account. Although the Bible was considered authoritative, the knowledge generated by science has been (...) so powerful and reliable in different things. While some people might use science to prove their own opinions, bringing an encroachment on the territory of religion, which theologians do not qualify as a scientific field. This paper has collected and analysed ideas from different scientists, philosophers, historians and theologians to examine the relationship between the Bible and Science, and the extent to which one is inductive to another. While an existence of a super intelligent designer was a common idea, confrontation focused on timeliness, creation story, divine action and miracles. Since religious people appeal to factual statements, while science is still developing and cannot prove the Bible or the existence of God with its lack of moral judgment, the Bible was qualified authoritative for faith and life. (shrink)

The theory of natural selection is a rich systematization of biological knowledge without a first principle. When formulations of a proposed principle of natural selection are examined carefully, each is seen to be exhaustively analyzable into a proposition about sources of fitness and a proposition about consequences of fitness. But whenever the fitness of an organic variety is well defined in a given biological situation, its sources are local contingencies together with the background of laws from disciplines other than the (...) theory of natural selection; and the consequences of fitness for the long range fate of organic varieties are essentially applications of probability theory. Hence there is no role and no need for a principle of the theory of natural selection, and any generalities that may hold in that theory are derivative rather than fundamental. (shrink)

This article identifies already existing theoretical and methodological commonalities between evolutionary biology and phenomenology, concentrating specifically on their common pursuit of origins. It identifies in passing theoretical support from evolutionary biology for present-day concerns in philosophy, singling out Sartre’s conception of fraternity as an example. It anchors its analysis of the common pursuit of origins in Husserl’s consistent recognition of the grounding significance of Nature and in his consistent recognition of animate forms of life other than human. It enumerates and (...) exemplifies five basic errors of continental philosophers with respect to Nature, errors testifying to a philosophical fundamentalism that distorts the intricate interconnections and relationships of Nature in favor of a preferred knowledge rooted in ontological reductionism. It shows that to discover and appreciate the common ground, one must indeed study “the things themselves.”. (shrink)

An examination of animate from reveals corporeal archetypes that underlie both human sexual behavior and the reigning Western biological paradigm of human sexuality that reworks the archetypes to enforce female oppression. Viewed within the framework of present-day social constructionist theory and Western biology, I show how both social constructionist feminists who disavow biology and biologists who reduce human biology to anatomy forget evolution and thereby forego understandings essential to the political liberation of women.

By now, even the kid down the street must be familiar with the functionalist's response to type-identity physicalism. Mental kinds like pain, love, the belief that Madison sits on an isthmus, etc., are not identical to physical kinds because it's conceptually possible that entities physically distinct in kind from human beings experience pain, love, beliefs that Madison sits on an isthmus, etc. Type-identity physicalism, in short, is baselessly chauvinistic in its rejection of the possibility of nonhuman minds.

This article is based on a perspective on circulation of knowledge that allows the consideration of science as the result of the encounter between diverse communities. We tell a story that constantly changes places, scales, and cultures in order to stress the importance of networks as an alternative to the centre/periphery trope, which entangles world histories of science. The result is a picture much more complex and intertwined than the one suggested by these simplifying dichotomies. We focus on a case (...) study that illuminates the process of knowledge production in non-European spaces of modernity. The return of the Society of Jesus to the newly independent nation-states of Latin America is the point of departure to analyse the circulation of a specific scientific idea in Ecuador: Darwin’s theory of biological evolution through natural selection. The article follows the paths of three different knowledge makers whose encounters are seen as sites of knowledge production: a religious order, a Latin American... (shrink)

At one level, this paper is a lament and a warning. I lament biologists borrowing well-known terms and then drastically and awkwardly changing their meanings, and I warn about the mischief this does. Biology''s public image is at stake, as is its general usefulness. At another level, I attempt to clarify the misnamed concepts, beyond what has been achieved in recent philosophical writings. This helps to account for the mischief, and to see how it might be avoidable. But the most (...) important thing about the paper is that, at a third level, it is an argument against physicalism and materialism, especially those variants which deny the autonomy of organisms and the existence of intrinsic goods. Interpreting biology from the point of view of those denials leads to unsatisfactory and even bizarre results. (shrink)

This article discusses the contribution to evolutionary theory of Leonard Darwin, the eighth child of Charles Darwin. By analysing the correspondence Leonard Darwin maintained with Ronald Aylmer Fisher in conjunction with an assessment of his books and other written works between the 1910s and 1930s, this article argues for a more prominent role played by him than the previously recognised in the literature as an informal mentor of Fisher. The paper discusses Leonard’s efforts to amalgamate Mendelism with both Eugenics and (...) Darwinism in order for the first to base their policies on new scientific developments and to help the second in finding a target for natural selection. Without a formal qualification in biological sciences and as such mistrusted by some “formal” scientists, Leonard Darwin engaged with key themes of Darwinism such as mimicry, the role of mutations on speciation and the process of genetic variability, arriving at important conclusions concerning the usefulness of Mendelian genetics for his father’s theory. (shrink)