Professor Grene's work on Aristotle is considered under three headings: teleology, form, and reductionism. A picture of Aristotle's philosophy of biology is sketched which stresses three elements: the place of living activity in the teleological account of the development and nature of organic structures; the functional nature of Aristotelian form; and the autonomy of biology as a natural science with its own basic principles. These elements are aspects of Aristotle's approach to biology with which Professor Grene has expressed sympathy.

The concept of "fitness" is a notion of central importance to evolutionary theory. Yet the interpretation of this concept and its role in explanations of evolutionary phenomena have remained obscure. We provide a propensity interpretation of fitness, which we argue captures the intended reference of this term as it is used by evolutionary theorists. Using the propensity interpretation of fitness, we provide a Hempelian reconstruction of explanations of evolutionary phenomena, and we show why charges of circularity which have been levelled (...) against explanations in evolutionary theory are mistaken. Finally, we provide a definition of natural selection which follows from the propensity interpretation of fitness, and which handles all the types of selection discussed by biologists, thus improving on extant definitions. (shrink)

I defend a view of the species category, pluralistic realism, which is designed to do justice to the insights of many different groups of systematists. After arguing that species are sets and not individuals, I proceed to outline briefly some defects of the biological species concept. I draw the general moral that similar shortcomings arise for other popular views of the nature of species. These shortcomings arise because the legitimate interests of biology are diverse, and these diverse interests are reflected (...) in different legitimate approaches to the classification of organisms. In the final section, I show briefly how the pluralistic approach can help to illuminate some areas of biological and philosophical dispute. (shrink)

Comprehensive evaluation and study of man's theories and knowledge of genetical characteristics and the evolutionary processes.

In the Preface of Animal Species and Evolution (1963), I wrote that it was "an attempt to summarize and review critically what we know about the biology and genetics of animal species and their role in evolution." The result was a volume of XIV ...

Essentialism--understood as the doctrine that there are natural kinds--can be sustained with respect to the most fundamental physical entities of the world, as I elsewhere argue. In this paper I take up the question of the existence of natural kinds among complex structures built out of these elementary ones. I consider a number of objections to essentialism, in particular Locke's puzzle about the existence of borderline cases. A number of recent attempts to justify biological taxonomy are critically examined. I conclude (...) that theory partially justifies such taxonomies but supports only a weaker form of essentialism. (shrink)

This paper gives an account of evolutionary explanations in biology. Briefly, the explanations I am primarily concerned with are explanations of adaptations. These explanations are contrasted with other nonteleological evolutionary explanations. The distinction is made by distinguishing the different kinds of questions these different explanations serve to answer. The sense in which explanations of adaptations are teleological is spelled out.

One possible interpretation of the species concept is that specifies are natural kinds. Another species concept is that species are individuals whose parts are organisms. Philip Kitcher takes seriously both these ideas; he sees a role for the genealogical/historical conception and also for the one that is “purely qualitative”. I criticize his ideas here. I see the genealogical conception at work in biological discussion of species and it is presupposed by an active and inventive research program, but the natural kind (...) concept is not. I also criticize Kitcher’s criticisms against Hull’s position that species are individuals, including Kitcher’s idea that species are sets of organisms, his explanations of why “all swans are white” isn’t law-like, his discussion of the example of multiple origination. Finally, I argue that given the current evolutionary developments, pluralism is the "null hypothesis" that we should attempt to refute. (shrink)

Oaks exemplify problems with the reproductive species concept which motivate a reconsideration of the use and nature of species. Ecology is important in the reconsideration. The species level is usually overemphasized in evolutionary thought; selection acts on phenotypes and any mutualistic units. Standard definitions tend to inhibit free conceptual progress. Multispecies, sets of broadly sympatric species that exchange genes, may occur among animals as well as plants and may conceivably bridge kingdoms. This phenomenon can be adaptively important. There may be (...) taxa without species. The degree of modality of adaptive zones can be investigated empirically. (shrink)

Much if not most recent literature in philosophy of biology concerns the extent to which biological theories conform to what is known as the "received" philosophical view of scientific theories, a descendant of the logical-empiricist view of theories. But the received view currently faces a competitor--a very different view of theories known as the "semantic" view. It is argued here that the semantic view is more sensitive to the nature and limitations of evolutionary theory than is the received view. In (...) particular, the semantic view better accomodates the fact that evolutionary theory is bound to change as a result of the evolutionary process itself. This unusual feature of evolutionary theory provides a good reason for reconsidering the received view and paying close attention to the semantic view. (shrink)

The paper characterizes Darwin's theory, providing a synthesis of recent historical investigations in this area. Darwin's reading of Malthus led him to appreciate the importance of population pressures, and subsequently of natural selection, with the help of the wedge metaphor. But, in itself, natural selection did not furnish an adequate account of the origin of species, for which a principle of divergence was needed. Initially, Darwin attributed this to geographical isolation, but later, following his work on barnacles which underscored the (...) significance of variation, and arising from his work on botanical arithmetic, he supposed that diversity allowed more places to be occupied in a given region. So isolation was not regarded as essential. Large regions with intense competition, and with ample variation spread by blending, would facilitate speciation. The notion of place was different from niche, and it is questioned whether Darwin's views on ecology were as modern as is commonly supposed. Two notions of struggle are found in Darwin's theory; and three notions of variation. Criticisms of his theory led him to emphasize the importance of variation over a range of forms. Hence the theory was populational rather than typological. The theory required a Lamarckian notion of inheritable changes initiated by the environment as a source of variation. Also, Darwin deployed a use/habit theory; and the notion of sexual selection. Selection normally acted at the level of the individual, though kin selection was possible. Group selection was hinted at for man. Darwin's thinking (and also the exposition of his theory) was generally guided by the domestic-organism analogy, which satisfied his methodological requirement of a vera causa principle. (shrink)

This paper shows that species are individuals with respect to evolutionary theory in the sense that the laws of the theory deal with species as irreducible wholes rather than as sets of organisms. 'Species X' is an instantiation of a primitive term of the theory. I present a sketch of a proof that it cannot be defined within the theory as a set of organisms; the proof relies not on details of my axiomatization but rather on a generally accepted property (...) of speciation; hence the same argument should work for any axiomatization which captures this generally accepted property of speciation. (shrink)

In his "Discussion" (1984), Elliott Sober offers some criticisms of the view about species--pluralistic realism--advocated in my 1984. Sober's comments divide into three parts. He attempts to show that species are not sets; he responds to my critique of David Hull's thesis that species are individuals; and he offers some arguments for the claim that species are "chunks of the genealogical nexus." I consider each of these objections in turn, arguing that each of them fails. I attempt to use Sober's (...) insightful critique to explain and defend pluralistic realism more fully. (shrink)

Although it is possible to regard a species as a set with a special internal structure, it is preferable to regard a species as an individual precisely to emphasize this internal structure. It is necessary to recognize, moreover, that two organisms that are part of a single entity with respect to one process need not be part of a single entity with respect to another process. Furthermore, choosing to regard two entities (with respect to one process) as conspecific is not (...) to deny that there are two entities within this species. Thus, the systematist need not propose formal names for every entity he discovers, but the names he proposes ought to be as faithful to the relationship among the entities as is possible. (shrink)

Many of the objections to the sociobiological research program arise from putative peculiarities of human beings and human societies when many of them actually arise from the nature of hierarchically organized systems. The levels of selection problem is no easier to handle at the gene-organism interface than at the organism-society interface. The unity of the genotype is just as problematic as the cohesiveness of the gene pool. One distinction which helps to reduce confusion is between replicators and interactors. Implications of (...) various versions of evolutionary theory for human beings are set out. (shrink)

Interprets biological theory while looking at work done on genetics, systematics and selection.