Different directed acyclic graphs may be Markov equivalent in the sense that they entail the same conditional independence relations among the observed variables. Chickering provided a transformational characterization of Markov equivalence for DAGs, which is useful in deriving properties shared by Markov equivalent DAGs, and, with certain generalization, is needed to prove the asymptotic correctness of a search procedure over Markov equivalence classes, known as the GES algorithm. For DAG models with latent variables, maximal ancestral graphs provide a neat representation (...) that facilitates model search. However, no transformational characterization -- analogous to Chickering's -- of Markov equivalent MAGs is yet available. This paper establishes such a characterization for directed MAGs, which we expect will have similar uses as it does for DAGs. (shrink)