This paper supplements an earlier one (Wassermann 1978b). Its views aim to reinforce those of Lewontin and other prominent evolutionists, but differ significantly from the opinions of some philosophers of science, notably Popper (1957) and Olding (1978). A basic distinction is made between 'laws' and 'theories of mechanisms'. The 'Theory of Evolution' is not characterized by laws, but is viewed here as a hypertheory which explains classifiable evolutionary phenomena in terms of subordinate classifiable theories of 'evolution-specific mechanisms' (ESMs), each of (...) which could apply to a host of species. Adaptations could result from ESMs that are rooted in molecular complementarities. The status of optimization theories that aim to predict best adapted states of organisms or populations is also discussed. (shrink)

We distinguish dynamical and statistical interpretations of evolutionary theory. We argue that only the statistical interpretation preserves the presumed relation between natural selection and drift. On these grounds we claim that the dynamical conception of evolutionary theory as a theory of forces is mistaken. Selection and drift are not forces. Nor do selection and drift explanations appeal to the (sub-population-level) causes of population level change. Instead they explain by appeal to the statistical structure of populations. We briefly discuss the implications (...) of the statistical interpretation of selection for various debates within the philosophy of biologythe `explananda of selection' debate and the `units of selection' debate. (shrink)

Stuart Kauffman here presents a brilliant new paradigm for evolutionary biology, one that extends the basic concepts of Darwinian evolution to accommodate recent findings and perspectives from the fields of biology, physics, chemistry and mathematics. The book drives to the heart of the exciting debate on the origins of life and maintenance of order in complex biological systems. It focuses on the concept of self-organization: the spontaneous emergence of order widely observed throughout nature. Kauffman here argues that self-organization plays an (...) important role in the emergence of life itself and may play as fundamental a role in shaping life's subsequent evolution as does the Darwinian process of natural selection. Yet until now no systematic effort has been made to incorporate the concept of self-organization into evolutionary theory. The construction requirements which permit complex systems to adapt remain poorly understood, as is the extent to which selection itself can yield systems able to adapt more successfully. This book explores these themes. It shows how complex systems, contrary to expectations, can spontaneously exhibit stunning degrees of order, and how this order, in turn, is essential for understanding the emergence and development of life on Earth. Topics include the new biotechnology of applied molecular evolution, with its important implications for developing new drugs and vaccines; the balance between order and chaos observed in many naturally occurring systems; new insights concerning the predictive power of statistical mechanics in biology; and other major issues. Indeed, the approaches investigated here may prove to be the new center around which biologicalscience itself will evolve. The work is written for all those interested in the cutting edge of research in the life sciences. (shrink)

Comprehensive evaluation and study of man's theories and knowledge of genetical characteristics and the evolutionary processes.

NOW IN PAPERBACK"€"Starting from a collection of simple computer experiments"€"illustrated in the book by striking computer graphics"€"Stephen Wolfram shows how their unexpected results force a whole new way of looking at the operation of our universe.

In this important book, Ellery Eells explores and refines philosophical conceptions of probabilistic causality. In a probabilistic theory of causation, causes increase the probability of their effects rather than necessitate their effects in the ways traditional deterministic theories have specified. Philosophical interest in this subject arises from attempts to understand population sciences as well as indeterminism in physics. Taking into account issues involving spurious correlation, probabilistic causal interaction, disjunctive causal factors, and temporal ideas, Professor Eells advances the analysis of what (...) it is for one factor to be a positive causal factor for another. A salient feature of the book is a theory of token level probabilistic causation in which the evolution of the probability of a later event from an earlier event is central. This will be a book of crucial significance to philosophers of science and metaphysicians; it will also prove stimulating to many economists, psychologists, and physicists. (shrink)

Through his S–R model of statistical relevance, Wesley Salmon offers a solution to the scientific explanation of objectively improbable events.

FEW HEROES LOWER their sights in the prime of their lives; triumph leads inexorably on, often to destruction. Alexander wept because he had no new worlds to conquer; Napoleon, overextended, sealed his doom in the depth of a Russian winter. But Charles Darwin did not follow the Origin of Species (1859) with a general defense of natural selection or with its evident extension to human evolution (he waited until 1871 to publish The Descent of Man). Instead, he wrote his most (...) obscure work, a book entitled: On the Various Contrivances by Which British and Foreign Orchids Are Fertilized by Insects (1862). (shrink)

How do fitness and natural selection relate to other evolutionary factors like architectural constraint, mode of reproduction, and drift? In one way of thinking, drawn from Newtonian dynamics, fitness is one force driving evolutionary change and added to other factors. In another, drawn from statistical thermodynamics, it is a statistical trend that manifests itself in natural selection histories. It is argued that the first model is incoherent, the second appropriate; a hierarchical realization model is proposed as a basis for a (...) statistical treatment. It emerges that natural selection does not cause evolution; it just is evolution. The theory incorporates relations of statistical correlation, but not the kind of causation found in fundamental physical processes. (shrink)

“Probabilistic Causation” designates a group of theories that aim to characterize the relationship between cause and effect using the tools of probability theory. The central idea behind these theories is that causes change the probabilities of their effects. This article traces developments in probabilistic causation, including recent developments in causal modeling. A variety of issues within, and objections to, probabilistic theories of causation will also be discussed.

The neutral theory often is presented as a theory of "noise" or silent changes at an isolated "molecular level", relevant to marking the steady pace of divergence, but not to the origin of biological structure, function, or complexity. Nevertheless, precisely these issues can be addressed in neutral models, such as those elaborated here in regard to scrambled ciliate genes, gRNA-mediated RNA editing, the transition from self-splicing to spliceosomal splicing, and the retention of duplicate genes. All of these are instances of (...) a more general scheme of "constructive neutral evolution" that invokes biased variation, epistatic interactions, and excess capacities to account for a complex series of steps giving rise to novel structures or operations. The directional and constructive outcomes of these models are not due to neutral allele fixations per se, but to these other factors. Neutral models of this type may help to clarify the poorly understood role of non-selective factors in evolutionary innovation and directionality. (shrink)

This chapter surveys the philosophical problems raised by the two Darwinian claims of the existence of a Tree of a life, and the explanatory power of natural selection. It explores the specificity of explanations by natural selection, emphasizing the high context-dependency of any process of selection. Some consequences are drawn about the difficulty of those explanations to fit a nomological model of explanation, and the irreducibility of their historic-narrative dimension. The paper introduces to the debates about units of selection, stating (...) the compelling force of genic selectionnism but highlighting some critiques. Then it addresses the limits of selectionist explanations : the compared status of selection, drift and phylogenetic inertia are investigated, and the debates over adaptationism are presented, with the aim of defining the varieties of adaptationisms as research programs. In order to assess the scope of natural selection, the chapter addresses weak and strong challenges to the Synthetic theory of evolution, both from paleontology (punctuated equilibria, Gould’s contingency thesis) and evolutionary theory of development. We finally sketch some consequences of evolutionary theory concerning philosophical questions about human nature, on the basis of the hypothesis of the universality of selectionist explanations: this part deals mostly with epistemology and psychology. (shrink)

Some naturalistic theories of consciousness give an essential role to teleology.1 This teleology is said to arise due to natural selection. Thus it is claimed that only certain states, namely, those that have been selected for by evolutionary pro- cesses because they contribute to (or once contributed to) an organism’s fitness, are conscious states. These theories look as if they are assigning a creative role to natural selection. If a state is conscious only if it has been selected for, then (...) selec- tion appears to be able to create a new feature of states, namely, their conscious nature. Yet, intuitively, natural selection cannot create anything. Natural selec- tion chooses certain features that already exist and makes them more (or less) prevalent in a population, but it cannot bring features into existence itself. Natu- ral selection can select for conscious states, but it cannot create them. This con- clusion has recently been argued for by Steven Horst (1999). If it is right, then teleological theories of conscious states should be rejected. A state cannot become a conscious experience in virtue of having been selected for by evolu- tionary process. (shrink)

Chalmers and others have argued that physicalist microexplanation is incapable of solving the ‘hard problem’ of consciousness. This article examines whether evolutionary accounts of the mind, such as those developed by Millikan, Dretske and Flanagan, can add anything to make up for the possible short falls of more reductionist accounts. I argue that they cannot, because evolutionary accounts explain by appeal to a selectional history that only comes into the picture if consciousness can first arise due to spontaneous mutation in (...) some individual organism, and explaining this emergence of consciousness from DNA and embryology calls for precisely the kind of structurally-based supervenience account that Chalmers et al. have objected to. Not only does an evolutionary account not succeed where a reductionist account fails; the evolutionary account presupposes the possibility of a reductionist account. (shrink)

When philosophers defend epiphenomenalist doctrines, they often do so by way of a priori arguments. Here we suggest an empirical approach that is modeled on August Weismann’s experimental arguments against the inheritance of acquired characters. This conception of how epiphenomenalism ought to be developed helps clarify some mistakes in two recent epiphenomenalist positions – Jaegwon Kim’s (1993) arguments against mental causation, and the arguments developed by Walsh (2000), Walsh, Lewens, and Ariew (2002), and Matthen and Ariew (2002) that natural selection (...) and drift are not causes of evolution. A manipulationist account of causation (Woodward 2003) leads naturally to an account of how macro- and micro-causation are related and to an understanding of how epiphenomenalism at different levels of organization should be understood. (shrink)