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  1. Biological normativity: a new hope for naturalism?Walter Veit - 2021 - Medicine, Health Care and Philosophy 24 (2):291-301.
    Since Boorse [Philos Sci 44(4):542–573, 1977] published his paper “Health as a theoretical concept” one of the most lively debates within philosophy of medicine has been on the question of whether health and disease are in some sense ‘objective’ and ‘value-free’ or ‘subjective’ and ‘value-laden’. Due to the apparent ‘failure’ of pure naturalist, constructivist, or normativist accounts, much in the recent literature has appealed to more conciliatory approaches or so-called ‘hybrid accounts’ of health and disease. A recent paper by Matthewson (...)
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  • Teleological theories of mental content.Peter Schulte & Karen Neander - 2022 - Stanford Encyclopedia of Philosophy.
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  • Developmental Systems and Evolutionary Explanation.P. E. Griffiths & R. D. Gray - 1994 - Journal of Philosophy 91 (6):277-304.
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  • An organizational account of biological functions.Matteo Mossio, Cristian Saborido & Alvaro Moreno - 2009 - British Journal for the Philosophy of Science 60 (4):813-841.
    In this paper, we develop an organizational account that defines biological functions as causal relations subject to closure in living systems, interpreted as the most typical example of organizationally closed and differentiated self-maintaining systems. We argue that this account adequately grounds the teleological and normative dimensions of functions in the current organization of a system, insofar as it provides an explanation for the existence of the function bearer and, at the same time, identifies in a non-arbitrary way the norms that (...)
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  • A Generalized Selected Effects Theory of Function.Justin Garson - 2017 - Philosophy of Science 84 (3):523-543.
    I present and defend the generalized selected effects theory (GSE) of function. According to GSE, the function of a trait consists in the activity that contributed to its bearer’s differential reproduction, or differential retention, within a population. Unlike the traditional selected effects (SE) theory, it does not require that the functional trait helped its bearer reproduce; differential retention is enough. Although the core theory has been presented previously, I go significantly beyond those presentations by providing a new argument for GSE (...)
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  • Evolution, Dysfunction, and Disease: A Reappraisal.Paul E. Griffiths & John Matthewson - 2018 - British Journal for the Philosophy of Science 69 (2):301-327.
    Some ‘naturalist’ accounts of disease employ a biostatistical account of dysfunction, whilst others use a ‘selected effect’ account. Several recent authors have argued that the biostatistical account offers the best hope for a naturalist account of disease. We show that the selected effect account survives the criticisms levelled by these authors relatively unscathed, and has significant advantages over the BST. Moreover, unlike the BST, it has a strong theoretical rationale and can provide substantive reasons to decide difficult cases. This is (...)
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  • Gender Is a Natural Kind with a Historical Essence.Theodore Bach - 2012 - Ethics 122 (2):231-272.
    Traditional debate on the metaphysics of gender has been a contrast of essentialist and social-constructionist positions. The standard reaction to this opposition is that neither position alone has the theoretical resources required to satisfy an equitable politics. This has caused a number of theorists to suggest ways in which gender is unified on the basis of social rather than biological characteristics but is “real” or “objective” nonetheless – a position I term social objectivism. This essay begins by making explicit the (...)
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  • Teleosemantics without etiology.Bence Nanay - 2014 - Philosophy of Science 81 (5):798-810.
    The aim of teleosemantics is to give a scientifically respectable, or ‘naturalistic’ theory of mental content. In the debates surrounding the scope and merits of teleosemantics a lot has been said about the concept of indication (or carrying information). The aim of this paper is to focus on the other key concept of teleosemantics: biological function. It has been universally accepted in the teleosemantics literature that the account of biological function one should use to flesh out teleosemantics is that of (...)
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  • (1 other version)Assertion: A Function First Account.Christoph Kelp - 2018 - Noûs 52 (2):411-442.
    This paper aims to develop a novel account of the normativity of assertion. Its core thesis is that assertion has an etiological epistemic function, viz. to generate knowledge in hearers. In conjunction with a general account of etiological functions and their normative import, it is argued that an assertion is epistemically good if and only if it has the disposition to generate knowledge in hearers. In addition, reason is provided to believe that it makes sense to regulate the practice of (...)
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  • How to Be a Function Pluralist.Justin Garson - 2018 - British Journal for the Philosophy of Science 69 (4):1101-1122.
    I distinguish two forms of pluralism about biological functions, between-discipline pluralism and within-discipline pluralism. Between-discipline pluralism holds that different theories of function are appropriate for different subdisciplines of biology and psychology. I provide reasons for rejecting this view. Instead, I recommend within-discipline pluralism, which emphasizes the plurality of function concepts at play within any given subdiscipline of biology and psychology.
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  • Biological Criteria of Disease: Four Ways of Going Wrong.John Matthewson & Paul Edmund Griffiths - 2017 - Journal of Medicine and Philosophy 1 (4).
    We defend a view of the distinction between the normal and the pathological according to which that distinction has an objective, biological component. We accept that there is a normative component to the concept of disease, especially as applied to human beings. Nevertheless, an organism cannot be in a pathological state unless something has gone wrong for that organism from a purely biological point of view. Biology, we argue, recognises two sources of biological normativity, which jointly generate four “ways of (...)
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  • Etiological theories of function: A geographical survey.David J. Buller - 1998 - Biology and Philosophy 13 (4):505-527.
    Formulations of the essential commitment of the etiological theory of functions have varied significantly, with some individual authors' formulations even varying from one place to another. The logical geography of these various formulations is different from what is standardly assumed; for they are not stylistic variants of the same essential commitment, but stylistic variants of two non-equivalent versions of the etiological theory. I distinguish these “strong” and “weak” versions of the etiological theory (which differ with respect to the role of (...)
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  • The epistemic significance of collaborative research.K. Brad Wray - 2002 - Philosophy of Science 69 (1):150-168.
    I examine the epistemic import of collaborative research in science. I develop and defend a functional explanation for its growing importance. Collaborative research is becoming more popular in the natural sciences, and to a lesser degree in the social sciences, because contemporary research in these fields frequently requires access to abundant resources, for which there is great competition. Scientists involved in collaborative research have been very successful in accessing these resources, which has in turn enabled them to realize the epistemic (...)
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  • The function debate in philosophy.Arno Wouters - 2005 - Acta Biotheoretica 53 (2):123-151.
    This paper reviews the debate on the notion of biological function and on functional explanation as this takes place in philosophy. It describes the different perspectives, issues, intuitions, theories and arguments that have emerged. The author shows that the debate has been too heavily influenced by the concerns of a naturalistic philosophy of mind and argues that in order to improve our understanding of biology the attention should be shifted from the study of intuitions to the study of the actual (...)
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  • Artifacts and Their Functions.A. W. Eaton - 2020 - In Ivan Gaskell & Sarah Anne Carter (eds.), The Oxford Handbook of History and Material Culture. Oxford University Press.
    How do artifacts get their functions? It is typically thought that an artifact’s function depends on its maker’s intentions. This chapter argues that this common understanding is fatally flawed. Nor can artifact function be understood in terms of current uses or capacities. Instead, it proposes that we understand artifact function on the etiological model that Ruth Millikan and others have proposed for the biological realm. This model offers a robustly normative conception of function, but it does so naturalistically by employing (...)
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  • Fitness and function.D. M. Walsh - 1996 - British Journal for the Philosophy of Science 47 (4):553-574.
    According to historical theories of biological function, a trait's function is determined by natural selection in the past. I argue that, in addition to historical functions, ahistorical functions ought to be recognized. I propose a theory of biological function which accommodates both. The function of a trait is the way it contributes to fitness and fitness can only be determined relative to a selective regime. Therefore, the function of a trait can only be specified relative to a selective regime. Apart (...)
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  • Are Biological Traits Explained by Their ‘Selected Effect’ Functions?Joshua R. Christie, Carl Brusse, Pierrick Bourrat, Peter Takacs & Paul E. Griffiths - 2022 - Australasian Philosophical Review 6 (4):335-359.
    The selected effects or ‘etiological’ theory of Proper function is a naturalistic and realist account of biological teleology. It is used to analyse normativity in philosophy of language, philosophy of mind, philosophy of medicine, and elsewhere. The theory has been developed with a simple and intuitive view of natural selection. Traits are selected because of their positive effects on the fitness of the organisms that have them. These ‘selected effects’ are the Proper functions of the traits. Proponents argue that this (...)
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  • Function, selection, and construction in the brain.Justin Garson - 2012 - Synthese 189 (3):451-481.
    A common misunderstanding of the selected effects theory of function is that natural selection operating over an evolutionary time scale is the only functionbestowing process in the natural world. This construal of the selected effects theory conflicts with the existence and ubiquity of neurobiological functions that are evolutionary novel, such as structures underlying reading ability. This conflict has suggested to some that, while the selected effects theory may be relevant to some areas of evolutionary biology, its relevance to neuroscience is (...)
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  • Functional analysis and the species design.Karen Neander - 2017 - Synthese 194 (4).
    This paper argues that a minimal notion of function and a notion of normal-proper function are used in explaining how bodies and brains operate. Neither is Cummins’ notion, as originally defined, and yet his is often taken to be the clearly relevant notion for such an explanatory context. This paper also explains how adverting to normal-proper functions, even if these are selected functions, can play a significant scientific role in the operational explanations of complex systems that physiologists and neurophysiologists provide, (...)
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  • (1 other version)Four notions of biological function.Arno G. Wouters - 2003 - Studies in History and Philosophy of Science Part C: Studies in History and Philosophy of Biological and Biomedical Sciences 34 (4):633-668.
    I argue that there are at least four different ways in which the term ‘function’ is used in connection with the study of living organisms, namely: function as activity, function as biological role, function as biological advantage, and function as selected effect. Notion refers to what an item does by itself; refers to the contribution of an item or activity to a complex activity or capacity of an organism; refers to the value for the organism of an item having a (...)
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  • Parts and theories in compositional biology.Rasmus Grønfeldt Winther - 2006 - Biology and Philosophy 21 (4):471-499.
    I analyze the importance of parts in the style of biological theorizing that I call compositional biology. I do this by investigating various aspects, including partitioning frames and explanatory accounts, of the theoretical perspectives that fall under and are guided by compositional biology. I ground this general examination in a comparative analysis of three different disciplines with their associated compositional theoretical perspectives: comparative morphology, functional morphology, and developmental biology. I glean data for this analysis from canonical textbooks and defend the (...)
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  • The function of perceptual learning.Zoe Jenkin - 2023 - Philosophical Perspectives 37 (1):172-186.
    Our perceptual systems are not stagnant but can learn from experience. Why is this so? That is, what is the function of perceptual learning? I consider two answers to this question: The Offloading View, which says that the function of perceptual learning is to offload tasks from cognition onto perception, thereby freeing up cognitive resources (Connolly, 2019) and the Perceptual View, which says that the function of perceptual learning is to improve the functioning of perception. I argue that the Perceptual (...)
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  • Normal‐proper functions in the philosophy of mind.Andrew Rubner - 2022 - Philosophy Compass (7):1-11.
    This paper looks at the nature of normal-proper functions and the role they play in theories of representational content. More specifically: I lay down two desiderata for a theory which tries to capture what's distinctive of normal-proper functions and discuss two prominent theories which claim to satisfy them. I discuss the advantages of having normal-proper functions ground a theory of representational content. And, I look at both orthodox and heterodox versions of such theories.
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  • Computing Mechanisms Without Proper Functions.Joe Dewhurst - 2018 - Minds and Machines 28 (3):569-588.
    The aim of this paper is to begin developing a version of Gualtiero Piccinini’s mechanistic account of computation that does not need to appeal to any notion of proper (or teleological) functions. The motivation for doing so is a general concern about the role played by proper functions in Piccinini’s account, which will be evaluated in the first part of the paper. I will then propose a potential alternative approach, where computing mechanisms are understood in terms of Carl Craver’s perspectival (...)
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  • (1 other version)Four notions of biological function.Arno G. Wouters - 2002 - Studies in History and Philosophy of Science Part C: Studies in History and Philosophy of Biological and Biomedical Sciences 34 (4):633-668.
    I argue that there are at least four different ways in which the term ‘function’ is used in connection with the study of living organisms, namely: function as activity, function as biological role, function as biological advantage, and function as selected effect. Notion refers to what an item does by itself; refers to the contribution of an item or activity to a complex activity or capacity of an organism; refers to the value for the organism of an item having a (...)
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  • Representation in the genome and in other inheritance systems.Nicholas Shea - 2007 - Biology and Philosophy 22 (3):313-331.
    There is ongoing controversy as to whether the genome is a representing system. Although it is widely recognised that DNA carries information, both correlating with and coding for various outcomes, neither of these implies that the genome has semantic properties like correctness or satisfaction conditions, In the Scope of Logic, Methodology, and the Philosophy of Sciences, Vol. II. Kluwer, Dordrecht, pp. 387–400). Here a modified version of teleosemantics is applied to the genome to show that it does indeed have semantic (...)
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  • Self-re-production and functionality.Gerhard Schlosser - 1998 - Synthese 116 (3):303-354.
    Function and teleology can be naturalized either by reference to systems with a particular type of organization or by reference to a particular kind of history. As functions are generally ascribed to states or traits according to their current role and regardless of their origin, etiological accounts are inappropriate. Here, I offer a systems-theoretical interpretation as a new version of an organizational account of functionality, which is more comprehensive than traditional cybernetic views and provides explicit criteria for empirically testable function (...)
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  • A Taxonomy of Functions.Denis M. Walsh & André Ariew - 1996 - Canadian Journal of Philosophy 26 (4):493 - 514.
    There are two general approaches to characterising biological functions. One originates with Cummins. According to this approach, the function of a part of a system is just its causal contribution to some specified activity of the system. Call this the ‘C-function’ concept. The other approach ties the function of a trait to some aspect of its evolutionary significance. Call this the ‘E-function’ concept. According to the latter view, a trait's function is determined by the forces of natural selection. The C-function (...)
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  • Artifact.Risto Hilpinen - 1999 - Stanford Encyclopedia of Philosophy.
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  • (1 other version)A Framework for the Emotional Psychology of Group Membership.Taylor Davis & Daniel Kelly - 2021 - Review of Philosophy and Psychology:1-22.
    The vast literature on negative treatment of outgroups and favoritism toward ingroups provides many local insights but is largely fragmented, lacking an overarching framework that might provide a unified overview and guide conceptual integration. As a result, it remains unclear where different local perspectives conflict, how they may reinforce one another, and where they leave gaps in our knowledge of the phenomena. Our aim is to start constructing a framework to help remedy this situation. We first identify a few key (...)
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  • (1 other version)A Dual-Aspect Theory of Artifact Function.Marc Artiga - 2023 - Erkenntnis 88 (4):1533-1554.
    The goal of this essay is to put forward an original theory of artifact function, which takes on board the results of the debate on the notion of biological function and also accommodates the distinctive aspects of artifacts. More precisely, the paper develops and defends the Dual-Aspect Theory, which is a monist account according to which an artifact’s function depends on intentional and reproductive aspects. It is argued that this approach meets a set of theoretical and meta-theoretical desiderata and is (...)
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  • Trustworthy artificial intelligence.Mona Simion & Christoph Kelp - 2020 - Asian Journal of Philosophy 2 (1):1-12.
    This paper develops an account of trustworthy AI. Its central idea is that whether AIs are trustworthy is a matter of whether they live up to their function-based obligations. We argue that this account serves to advance the literature in a couple of important ways. First, it serves to provide a rationale for why a range of properties that are widely assumed in the scientific literature, as well as in policy, to be required of trustworthy AI, such as safety, justice, (...)
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  • Ascribing functions to technical artefacts: A challenge to etiological accounts of functions.Pieter E. Vermaas & Wybo Houkes - 2003 - British Journal for the Philosophy of Science 54 (2):261-289.
    The aim of this paper is to evaluate etiological accounts of functions for the domain of technical artefacts. Etiological theories ascribe functions to items on the basis of the causal histories of those items; they apply relatively straightforwardly to the biological domain, in which neo-Darwinian evolutionary theory provides a well-developed and generally accepted background for describing the causal histories of biological items. Yet there is no well-developed and generally accepted theory for describing the causal history of artefacts, so the application (...)
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  • I—Fundamental Powers, Evolved Powers, and Mental Powers.Alexander Bird - 2018 - Aristotelian Society Supplementary Volume 92 (1):247-275.
    Powers have in recent years become a central component of many philosophers’ ontology of properties. While I have argued that powers exist at the fundamental level of properties, many other theorists of powers hold that there are also non-fundamental powers. In this paper I articulate my reasons for being sceptical about the existing reasons for holding that there are non-fundamental powers. However, I also want to promote a different argument for the existence of a certain class of non-fundamental powers: properties (...)
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  • Do transposable elements have functions of their very own?Justin Garson - 2022 - Biology and Philosophy 37 (3):1-18.
    Philosophers who study the problem of biological function often begin their deliberations by reflecting on the functions of parts of animals, or the behavior of animals. Applying theories of biological function to unconventional or borderline cases can help us to better evaluate and refine those theories. This is the case when we consider whether parts of transposable elements —bits of “selfish” DNA that move about within a host genome—have functions of their own, that is, whether the parts of TEs have (...)
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  • The Function of Perception.Peter J. Graham - 2014 - In Abrol Fairweather (ed.), Virtue Scientia: Bridges between Virtue Epistemology and Philosophy of Science. Dordrecht, Netherlands: Synthese Library. pp. 13-31.
    What is the biological function of perception? I hold perception, especially visual perception in humans, has the biological function of accurately representing the environment. Tyler Burge argues this cannot be so in Origins of Objectivity (Oxford, 2010), for accuracy is a semantical relationship and not, as such, a practical matter. Burge also provides a supporting example. I rebut the argument and the example. Accuracy is sometimes also a practical matter if accuracy partly explains how perception contributes to survival and reproduction.
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  • Replicator II – judgement day.Paul E. Griffiths & Russell D. Gray - 1997 - Biology and Philosophy 12 (4):471-492.
    The Developmental Systems approach to evolution is defended against the alternative extended replicator approach of Sterelny, Smith and Dickison (1996). A precise definition is provided of the spatial and temporal boundaries of the life-cycle that DST claims is the unit of evolution. Pacé Sterelny et al., the extended replicator theory is not a bulwark against excessive holism. Everything which DST claims is replicated in evolution can be shown to be an extended replicator on Sterelny et al.s definition. Reasons are given (...)
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  • Function, persistence, and selection: Generalizing the selected-effect account of function adequately.Pierrick Bourrat - 2021 - Studies in History and Philosophy of Science Part A 90 (C):61-67.
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  • Biological function, adaptation, and natural design.Colin Allen & Marc Bekoff - 1995 - Philosophy of Science 62 (4):609-622.
    Recently something close to a consensus about the best way to naturalize the notion of biological function appears to be emerging. Nonetheless, teleological notions in biology remain controversial. In this paper we provide a naturalistic analysis for the notion of natural design. Many authors assume that natural design should be assimilated directly to function. Others find the notion problematic because it suggests that evolution is a directed process. We argue that both of these views are mistaken. Our naturalistic account does (...)
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  • Selected effects and causal role functions in the brain: the case for an etiological approach to neuroscience.Justin Garson - 2011 - Biology and Philosophy 26 (4):547-565.
    Despite the voluminous literature on biological functions produced over the last 40 years, few philosophers have studied the concept of function as it is used in neuroscience. Recently, Craver (forthcoming; also see Craver 2001) defended the causal role theory against the selected effects theory as the most appropriate theory of function for neuroscience. The following argues that though neuroscientists do study causal role functions, the scope of that theory is not as universal as claimed. Despite the strong prima facie superiority (...)
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  • Function in the Light of Frequency-dependent Selection.Samir Okasha - 2022 - Australasian Philosophical Review 6 (4):386-399.
    Christie, Brusse, et al. claim that the ‘selected effect’ (SE) theory of function is premised on a simplistic view of evolution. In complex evolutionary scenarios, in particular those involving frequency-dependent selection (FDS), the SE theory fails, they argue, since citing a trait’s SE function does not serve to explain why the trait exists. I argue that where FDS leads to a stable equilibrium, at which all individuals’ trait values constitute a ‘best response’ to the rest of the population, the SE (...)
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  • How objective are biological functions?Marcel Weber - 2017 - Synthese 194 (12):4741-4755.
    John Searle has argued that functions owe their existence to the value that we put into life and survival. In this paper, I will provide a critique of Searle’s argument concerning the ontology of functions. I rely on a standard analysis of functional predicates as relating not only a biological entity, an activity that constitutes the function of this entity and a type of system but also a goal state. A functional attribution without specification of such a goal state has (...)
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  • (1 other version)Teleological Notions in Biology.Colinn D. Allen - 2012 - In Ed Zalta (ed.), Stanford Encyclopedia of Philosophy. Stanford, CA: Stanford Encyclopedia of Philosophy.
    Teleological terms such as "function" and "design" appear frequently in the biological sciences. Examples of teleological claims include: A (biological) function of stotting by antelopes is to communicate to predators that they have been detected. Eagles' wings are (naturally) designed for soaring. Teleological notions were commonly associated with the pre-Darwinian view that the biological realm provides evidence of conscious design by a supernatural creator. Even after creationist viewpoints were rejected by most biologists there remained various grounds for concern about the (...)
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  • Biological functions and natural selection: a reappraisal.Marc Artiga - 2021 - European Journal for Philosophy of Science 11 (2):1-22.
    The goal of this essay is to assess the Selected-Effects Etiological Theory of biological function, according to which a trait has a function F if and only if it has been selected for F. First, I argue that this approach should be understood as describing the paradigm case of functions, rather than as establishing necessary and sufficient conditions for function possession. I contend that, interpreted in this way, the selected-effects approach can explain two central properties of functions and can satisfactorily (...)
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  • What are Functions Good For?Justin Garson - 2022 - Australasian Philosophical Review 6 (4):374-385.
    Christie, Brusse, et al. argue that the selected effects theory of function (SE) doesn’t do what it’s supposed to do: namely, show how functions can be explanatory. They survey some well-known evolutionary dynamics such as arms races, frequency-dependent fitness, and environmental heterogeneity, some of which have been discussed in the functions literature for decades. They argue that SE only seems to work because SE theorists ignore these dynamics. Their argument fails because they misrepresent what functions are supposed to explain and (...)
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  • Reasoning Is for Arguing: Understanding the Successes and Failures of Deliberation.Hugo Mercier & Hélène Landemore - unknown
    Theoreticians of deliberative democracy have sometimes found it hard to relate to the seemingly contradictory experimental results produced by psychologists and political scientists. We suggest that this problem may be alleviated by inserting a layer of psychological theory between the empirical results and the normative political theory. In particular, we expose the argumentative theory of reasoning that makes the observed pattern of findings more coherent. According to this theory, individual reasoning mechanisms work best when used to produce and evaluate arguments (...)
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  • Does proper function come in degrees?John Matthewson - 2020 - Biology and Philosophy 35 (4):1-18.
    Natural selection comes in degrees. Some biological traits are subjected to stronger selective force than others, selection on particular traits waxes and wanes over time, and some groups can only undergo an attenuated kind of selective process. This has downstream consequences for any notions that are standardly treated as binary but depend on natural selection. For instance, the proper function of a biological structure can be defined as what caused that structure to be retained by natural selection in the past. (...)
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  • (1 other version)A Framework for the Emotional Psychology of Group Membership.Taylor Davis & Daniel Kelly - 2024 - Review of Philosophy and Psychology 15 (2):333-354.
    The vast literature on negative treatment of outgroups and favoritism toward ingroups provides many local insights but is largely fragmented, lacking an overarching framework that might provide a unified overview and guide conceptual integration. As a result, it remains unclear where different local perspectives conflict, how they may reinforce one another, and where they leave gaps in our knowledge of the phenomena. Our aim is to start constructing a framework to help remedy this situation. We first identify a few key (...)
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  • Junk or functional DNA? ENCODE and the function controversy.Pierre-Luc Germain, Emanuele Ratti & Federico Boem - 2014 - Biology and Philosophy 29 (6):807-831.
    In its last round of publications in September 2012, the Encyclopedia Of DNA Elements (ENCODE) assigned a biochemical function to most of the human genome, which was taken up by the media as meaning the end of ‘Junk DNA’. This provoked a heated reaction from evolutionary biologists, who among other things claimed that ENCODE adopted a wrong and much too inclusive notion of function, making its dismissal of junk DNA merely rhetorical. We argue that this criticism rests on misunderstandings concerning (...)
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  • In What Sense Does ‘Nothing Make Sense Except in the Light of Evolution’?Paul Edmund Griffiths - 2009 - Acta Biotheoretica 57 (1-2):11-32.
    Dobzhansky argued that biology only makes sense if life on earth has a shared history. But his dictum is often reinterpreted to mean that biology only makes sense in the light of adaptation. Some philosophers of science have argued in this spirit that all work in ‘proximal’ biosciences such as anatomy, physiology and molecular biology must be framed, at least implicitly, by the selection histories of the organisms under study. Others have denied this and have proposed non-evolutionary ways in which (...)
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